Ramon A. Carreno

Professor of Zoology


B.S., University of Toronto
M.S., Lakehead University
Ph.D., University of Guelph


Ramon Carreno’s most recent research has focused on the systematics of parasitic nematodes (worms) in mammals, based on a project funded by the National Science Foundation. Other research interests include diagnostic parasitology, as well as genotyping and systematics of other complex parasitic organisms.

Research approaches incorporate light and electron microscopy, as well as cloning and DNA sequencing. He is the author of more than 40 articles, reports, and book chapters.

He also serves as Ohio Wesleyan’s pre-veterinary medicine program adviser. In addition to English, Carreno is fluent in French, German, and Spanish. Read his complete CV.

Areas of Interest / Expertise

  • The evolution of lung worms
  • Parasitic nematodes
  • Diagnostic parasitology

Publications / Presentations

  • Carreno, Ramon A., et al. (2008). “Equinurbia blakei n. sp. (Nematoda: Strongyloidea: Strongylidae) from the African Forest Elephant, Loxodonta cyclotis.” Comparative Parasitology, 75: 196–199.
  • Carreno, Ramon A., et al. (2008). “Dispharynx, Echinuria, and Streptocara.” Parasitic Diseases of Wild Birds. Ed. Carter Atkinson, et al. Ames, Iowa: Wiley-Blackwell: 326–342.
  • Carreno, Ramon A., et al. (2009). “Mermithid Nematode Infection in a Colony of Blue-Winged Grasshoppers (Tropidacris Collaris).” Journal of Zoo and Wildlife Medicine, 39: 488–492.
  • Carreno, Ramon A., et al. (2008). “A New Species of Paraheligmonella (Nematoda, Heligmonellidae), Parasite of Sylvilagus Floridanus (Leporidae) from Costa Rica.” Revista Mexicana de Biodiversidad, 79: 121S–128S.
  • Nadler, S.A., Carreno, R.A., Mejia-Madrid, H., Ullberg, J., Pagan, C., Houston, R., and Hugot, J.P. (2007). Molecular phylogeny of clade III nematodes reveals multiple origins of tissue parasitism. Parasitology. In Press.
  • Carreno, R.A. (2007). Description of a new species of Thelastoma Leidy, 1849 (Nematoda: Oxyurida: Thelastomatidae) from the millipede Euryurus leachii (Gray, 1832) in Ohio, U.S.A. Comparative Parasitology. In Press.
  • Conrad, P.A., A.M. Kjemtrup, R.A. Carreno, J. Thomford, K. Wainwright, M. Eberhard, R. Quick, S.R. Telford III, and B.L. Herwaldt. (2006). Description of Babesia duncani n.sp. (Apicomplexa: Babesiidae) from humans and its differentiation from other piroplasms. International Journal for Parasitology, 36: 779–789.
  • Kjemtrup, A.M., K. Wainwright, M. Miller, B.L. Penzhorn, and R.A. Carreno. (2006). Babesia conradae, sp. nov., a small canine Babesia identified in California. Veterinary Parasitology, 138: 103–111.

Contact Info


Schimmel/Conrades Science Center #344
Ohio Wesleyan University
Delaware, OH 43015
P 740-368-3893
E racarren@owu.edu