Ramon A. Carreno

Professor of Biological Sciences


  • B.S., University of Toronto
  • M.S., Lakehead University
  • Ph.D., University of Guelph


Ramon Carreno’s most recent research has focused on the systematics of parasitic nematodes in mammals, birds, and arthropods, as part of a broader interest in wildlife parasitology and wildlife diseases.

Research approaches incorporate light and electron microscopy, as well as cloning and DNA sequencing.

Dr. Carreno also serves as Ohio Wesleyan’s pre-veterinary medicine program adviser. In addition to English, Carreno is fluent in French, German, and Spanish.

Areas of Interest / Expertise

  • parasite taxonomy
  • wildlife diseases
  • diagnostic parasitology

Recent Publications / Presentations

  • Carreno, R. A. 2017. New species of Hammerschmidtiella Chitwood, 1932, and Blattophila Cobb, 1920, and new geographical records for Severianoia annamensis Van Luc & Spiridonov, 1993 (Nematoda: Oxyurida: Thelastomatoidea) from cockroaches (Insecta: Blattaria) in Ohio and Florida, U.S.A. Zootaxa 4226: 429-441.
  • Sinnott, D., R. A. Carreno, and Henri Herrera. 2015. Distribution of thelastomatoid nematodes (Nematoda: Oxyurida) in endemic and introduced cockroaches on the Galápagos Island archipelago, Ecuador. Journal of Parasitology 101: 445-457.
  • Carreno, R. A. 2014. The systematics and evolution of pinworms (Nematoda: Oxyurida: Thelastomatoidea) from invertebrates. Journal of Parasitology 100: 553-560.
  • Kim, T., J. Kim, S. Cho, G. Min, C. Park, R. A. Carreno, S. A. Nadler, and J. Park. 2014. Phylogenetic implications for Rhigonematomorpha based on the complete mitochondrial genome of Rhigonema thysanophora (Nematoda: Chromadorea). Zoologica Scripta 43: 289-303.
  • Hidalgo, M. R., S., A. Martinez, R. A. Carreno, S. Gonzalez M. C. Ferreras, and N. Diez. 2014. Levels of infection, pathology, and nodule size of Onchocerca flexuosa (Nematoda: Onchocercidae) in red deer Cervus elaphus from northern Spain. Journal of Helminthology. doi: 10.1017/S0022149X1400011X (print: 2015, vol. 89: 326-334)
  • Jimenez, F. A., R. A. Carreno, and S. L. Gardner. 2013. Aspidodera kinsellai, n.sp. (Nematoda: Heterakoidea) from nine-banded armadillos in Middle America with notes on phylogeny and host-parasite biogeography. Journal of Parasitology 99: 1056-1061.
  • Carreno, R. A., D. Ordosch, J. K. Koltek, D. R. Hamill, and L. Tuhela. 2013. First United States Records of the rhigonematid genera Heth and Ruizia (Nematoda: Rhigonematida) from the introduced millipede, Anadenobolus monilicornis (Diplopoda: Rhinocricidae) in Key Largo, Florida. Comparative Parasitology 80: 225-232.
  • Carreno, R. A., D. Caporossi, M. S. Beade, C. A. Marull, M. M. Uhart, D. D. Markwardt, and S. A. Nadler. 2012. Discovery of an undescribed protostrongylid nematode from the endangered pampas deer (Ozotoceros bezoarticus celer) in Argentina. Journal of Wildlife Diseases 48: 724-731.
  • Matsubayashi, M., R. A. Carreno, H. Tani, R. Yoshiuchi, T. Kanai, I. Kimata, S. Uni, M. Furuya, and K. Sasai. 2011. Phylogenetic identification of Cystoisospora spp. from dogs, cats, and raccoon dogs in Japan. Veterinary Parasitology 176: 270-274.
  • Carreno, R. A., and L. Tuhela. 2011. Thelastomatid nematodes (Oxyurida: Thelastomatoidea) from the peppered cockroach, Archimandrita tesselata (Insecta: Blattaria) in Costa Rica. Comparative Parasitology 78: 39-55.
  • Carreno, R. A., N. Diez-Baños, M. Hidalgo-Argüello, and S. A. Nadler. 2009. Characterization of Dictyocaulus species (Nematoda: Trichostrongyloidea) from three species of wild ruminants in northwestern Spain. Journal of Parasitology 95: 966-970.

Conference Presentations

  • Chu, R.A. Carreno, S. Koba, K.M. Davis, and C.M. Nagamine. 2014. Identification of and Surveillance for Syphacia Pinworms in a Colony of Golden Hamsters (Mesocricetus auratus). 65th National Meeting of the American Association for Laboratory Animal Science. San Antonio, Texas, October 19-23, 2014. Poster.
  • Sinnott, D. M., R. A. Carreno, and H. W. Herrera. 2014. Distribution of thelastomatoid nematodes (Oxyurida: Thelastomatoidea) in endemic and introduced cockroaches from the Galápagos Islands, Ecuador. The 89th Annual Meeting of the American Society of Parasitologists. New Orleans, Louisiana, July 24-27, 2014. Presented by D. Sinnott, OWU class of 2013.
  • Hidalgo, M.R., S. González, A. Martínez, R. A. Carreño, M. C. Ferreras, and N. Diez. 2013. Histological and morphological characteristics of Onchocerca flexuosa in red deer. Congreso de la Sociedad Española de Parasitologia. Gran Canaria 17-20 September, 2013. Poster.
  • Carreño, R.A. A. Martínez, D. D. Markwardt, N. Diez, and M. R. Hidalgo. 2013. Trichuris spp. from red deer (Cervus elaphus) in northern Spain. Congreso de la Sociedad Española de Parasitologia. Gran Canaria 17-20 September, 2013. Poster.
  • Oxyurid nematodes from cockroaches (Blattaria) in southern Florida with revised morphological characters for Euryconema paradisa and Protrelloides paradoxa. J.K. Koltek, R.A. Carreno, D.M. Ordosch, D.R. Hamill (2012). Poster presented at the 87th meeting of the American Society of Parasitologists, Richmond, Virginia, July 13-16, 2012. Presented by Josie Koltek, OWU Class of 2012.
  • Endoparasite survey in bobcats (Lynx rufus rufus) from Ohio. Markah Frost, Sarah Johnston, Suzie Prange, and Ramon A. Carreno. Poster. The 63st Annual Midwestern Conference of Parasitologists. Saint Mary’s College, Notre Dame, Indiana June 23-25, 2011.

Contact Info


Schimmel/Conrades Science Center #344
Ohio Wesleyan University
Delaware, OH 43015
P 740-368-3893
E racarren@owu.edu